In other words the tx fees are paid in iota tokens? If not what SPECIFIC action individuals do is driving demand for IOTA tokens?
Iota will enable nano payment and an efficient free market for IoT devices to trade data. In the IoT data trading marketplace, Iota tokens are the medium to trade data just like the Petrodollars in the international oil marketplace.
I think your next natural question will be "why is it possible?". It is possible because Iota has the potential to create the first IoT marketplace and companies and devices will have to use it. It is why Bitcoin is worthy much more than any other coins, first mover advantage.