The miners!
Honestly, the high fees of Bitcoin is not really good for anyone except the miners.
I would say it is beneficial for Bitcoin miners and Bitcoin mining pools. Nowadays, chance to find a Bitcoin block as a solo miner is very low, hence miners have to join Bitcoin mining pools and they will have to share part of their rewards (block subsidy and transaction fee) to mining pool operators.
It is like from 0.5 % to 4%. [1]
[1]
https://en.bitcoin.it/wiki/Comparison_of_mining_pools