^^^ But the question is. are animal feelings only complex computer (brain) responses? Or do they somehow have emotions. The OP tests suggest that there is far less emotion than owners might think.

But, in humans, feelings are not brain responses? every emotion is produced by a brain. The only difference between us and dogs is that we are aware of this emotion and are able to express it verbally.
Do you think that a shrimp, a fish or a mosquito feel emotions? I don't think so. But I think a dog does have emotions, because it shows them quite clearly.